I am an evolutionary biologist and work on the evolution of drug resistance in HIV. I want to understand what determines the rate of evolution of drug resistance, so that we can find ways to halt the evolution of drug resistance.

Before switching to HIV, I worked on ants, sympatric speciation and soft sweeps. I also write and tweet and make video’s.

If you have questions or would like to discuss an idea, feel free to drop me a line (pennings at!

My complete CV is here: Pennings_CV_2013_1 

Mini CV

Since September 2014: assistant professor in the Biology Department of San Francisco State University.

November 2012 – August 2014: postdoc with Dmitri Petrov at Stanford, project partly funded by HFSP.

June 2010 -Oct 2012 : postdoc with John Wakeley in the OEB department at Harvard, working on HIV drug resistance, project funded by HFSP.

2009 – 2010: postdoc with Susanne Foitzik on ant population genetics project funded by the German Science Foundation.

2006 – 2009: coordinator of the “Munich Graduate Program for Evolution, Ecology and Systematics” funded by VolkswagenFoundation.

January 2007: defended PhD thesis titled “Models of Adaptation and Speciation” (awarded highest grade: summa cum laude).

2003 – 2006: PhD in Joachim Hermisson‘s group at the University of Munich (LMU).

1999 – 2003: worked in the field of science education and started own company “De Praktijk”.

1994: switched to University of Amsterdam to study biology.

1993: started undergraduate studies in Aberdeen, Scotland.

1975 – 1993: grew up in Castricum on the coast of The Netherlands.


One Response to “Bio”

  1. David Stern (@sterndavidi) July 29, 2018 at 3:26 pm #

    I agree that some of those comments seem to be sexist. Having a foreign PhD also makes things tough. But I don’t think the comments about contribution are neccessarily sexist. I have written similar comments about men, myself. People do look for first author papers and in fields like economics single author papers. People do look for evidence of collaboration with people other than the advisor. Now I just looked at your track record on Scopus and I see there are three papers on “soft sweeps” and two have your name first, though the first and most cited has your advisor’s name first, which is unfortunate. I think this reviewer just looked at that and didn’t notice that the other two parts have your name first and came to the wrong conclusion. So that is bad.

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